Equine Physiology Workbook

Mechanical Digestion in the Small Intestine

 2 types of movements: Segmentations and Migrating Motility Complexes (a type of peristalsis). Both are mainly governed by the myenteric plexus.  Segmentations : localized mixing contractions that occur in portions of the intestine distended by a large volume of chyme. Segmentations mix chyme with digestive juices and bring particles of food in contact with the mucosa for absorption; they do not push contents along the tract. Essentially, the chyme sloshes back and forth.  Migrating Motility Complex (MMC): begins in the lower portion of the stomach and pushes chyme forward along a short stretch of the small intestine before dying out. The MMC slowly migrates down the small intestine reaching the end of the ileum in 90-120 minutes. Another MMC then begins in the stomach. All totaled, chyme remains in the small intestine for 45 min -3 hours, depending on the diet.

Chemical Digestion in the Small Intestine

 Chyme entering the small intestine contains partially digested carbohydrates, proteins and lipids. The completion of the digestion of these is a collective effort of pancreatic juice, bile, and intestinal juice in the small intestine.

The digestion of carbohydrates occurs with the following enzymes:

 Amylase – breaks starch down to simple sugars  α – Dextrinase, Sucrase, Lactase, Maltase (all brush-border enzymes) assist with starch breakdown  Digestion of carbohydrates ends with the production of monosaccharides, which the digestive system is able to absorb.  Carbohydrates exist in the equine diet in several forms:  Simple sugars  mono and disaccharides – found in grains, molasses, fruit, some in grasses  carbohydrates  polysaccharides  Starch in hay and grain  Fibre (branched chains of glucose units) in hay and grain.

 In the horse, the main dietary components are starch and fibre (cellulose)

Digestion of Proteins occurs with the following enzymes:

 Proteases such as Pepsin, enzymes of pancreatic juice and Peptidases (brush-border enzymes) break proteins into amino acids.

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